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GEN's Cryptogram Challenge: ELISA Redux proved to be very difficult to solve. It took 11 weeks to crack! Here are all the clues we revealed since the start of the contest.

Clue 1: The solution is a unique verbal statement. The color shown is not trivial.
(This clue is linked to Clue 5.)

Clue 2: Only letters are specifically coded. Color transparency is important to the solution.
(Blanks are not specifically coded. Specific color transparencies encode specific letters. Also see Clue 5 for assistance with allowed grid positions within which letters may be encoded.)

Clue 3: Letters are separated from one another by arbitrary numbers of blank grid positions.
(There are variable numbers of blanks between not only words but also the letters within words.)

Clue 4: The puzzle cannot be solved by letter-frequency analysis.
(The letters in the statement are used only once.)

Clue 5: The color spectrum ROYGBIV can be considered to be numeric. The row designations A–H may be assigned numbers 1–8; a special relationship exists between them and the column numbers as reflected within the cells themselves.
(The color green is the fourth in this series; see Clue 7. Row and column numbers are multiplied in order to provide a unique number for each cell.)

Clue 6: A special relationship between row number, column number, and color spectral position determines the allowable locations within which cells can be assigned letters, but not all allowed positions are filled with letters.
(See Clue 7.)

Clue 7: The multiple of row and column designations for each cell are divided by the color number in the spectrum ROYGBIV. In some cells, this results in an integer value. Only in these cells can letters be encoded.

Clue 8: Only even numbered transparencies are allowed.

Clue 9: Reading is on a diagonal.
(The answer reads from lower left to upper right within those cells that are allowed to be encoded.)

Clue 10: The answer is four words long. There is one two-letter word, two three-letter words, and one four-letter word.

Clue 11: Only transparencies between 4 and 54 encode for letters.

Decode the cipher hidden in this 96-well plate.

GEN and Scintellix have formed a partnership to explore the hidden meaning found in biological imagery. The Cryptogram Challenge: ELISA Redux is the third installment of the partnership.